The latter is associated with modifications in modes of colony foundation. Tree branches and tree background are colored for queen morphology and founding behavior respectively, according to the parsimony ancestral reconstruction. Typical queen-worker dimorphism shown to the right to illustrate ratio T1/T2 (not to scale). Species with wingless queens are marked with an asterisk. Phylogeny was pruned from Moreau et al. (2006). Placement of Sphecomyrma† and Martialis after Grimaldi et al. (1997) and Rabeling et al. (2008), respectively. Metapolybia and Scolia wasps are included as outgroups. Data on the species are analyzed, and their morphology and type of colony founding behavior are summarized in Table 3. Numbers correspond to major taxonomic groups within Formicidae after Ward (2007): 1, Sphecomyrminae†; 2, Leptanillinae; 3, Martialinae; 4, Proceratiinae; 5, Amblyoponinae; 6, Paraponerinae; 7, Agroecomyrmecinae; 8, Ponerinae; 9, dorylomorphs; 10, myrmeciomorphs; 11, dolichoderomorphs; 12, ectaheteromorphs; 13, Formicinae; 14, Myrmicinae.